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Systematics of fishes, zoogeography of northwest fishes, endangered species recovery, early life history ecology of fishes, and ichthyology.
Ph.D. - 1976 - Marine Science, College of William and Mary, Gloucester Pt.M.S. - 1972 - Marine Science, College of William and Mary, Gloucester Pt.B.S. - 1969 - Cornell University, Ithaca